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In addition to pulling and anchoring, we also quantified other meaningful aspects of individual and collective movement, including individuals’ speed, tortuosity of movement (low tortuosity equating to highly directed motion), distance to the group centre (centroid), the relative angle between the heading direction of the initiator and follower, and the relative angle between the individual’s heading direction and its direction to the group centroid, as well as group-level properties, such as the mean distance of individuals to the group’s centroid (that is, the spread of the group) as well as kinematics of the group’s centroid, including displacement (travelled distance), speed and tortuosity (details in Methods)
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